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Marley Fox, orphan, foundling, and poacher, is born towards the end of the 19th century at a time when humans are under attack from the Dark Angels who harvest souls for their own nefarious aims. The youngster's only family is Black Annie, a recluse who has occult powers. The Angels of Light are attempting to protect humans, and Marley is to play a part in the fight against those who would destroy their world. While out poaching, Marley is shot and badly injured. During his fight for survival, Marley has to battle a vicious Ice Angel. He is thrown in prison, where he is befriended by a huge Irishman, James Finnegan. An attempt is made to kill Marley. Finnegan and Marley escape and go on the run through England. Eventually, they take refuge in Ireland. They are still not safe, and they are attacked by the Dark Forces. It is one crisis after another as Marley and James battle to survive. The future fortunes of the world hinge on the ability of Marley and James to survive against the forces of the Angels of Darkness.

Having survived the best efforts of the Dark Angels to kill him, Marley Fox is conscripted into the British army along with his companion, James Finnegan. The two are sent to South Africa to fight in the war against the Boers, the brutality of which swallows them up as they are thrown into battle after battle. In the midst of the war, the Dark Angels send supernatural forces against them, and old enemies from England suddenly appear and do their best to finish off the pair. As enemies old and new close in on them, their survival is very much in doubt. Only their courage and fighting skills against seemingly insurmountable odds keep the friends going. Will they manage to stay alive as the attacks on them increase in intensity?